Can LoRA be applied to different base models?

[CJ0109]

I've always been curious about how LoRA works. For example, if I trained my LoRA parameters on SD-1-4, can I apply it directly to the base model of SD-1-5? According to my previous knowledge, this should not work, because this is equivalent to changing the model parameters of SD-1-5, which will cause the inference to fail.
But that doesn't seem to be the case. When I use Chilloutmix, different LoRA models such as JapaneseDoll, KoreaDoll can be combined into the base model with different weights, and it works quite well.